Fig. 2

Conchological and anatomical comparison of Pseudosuccinea columella (top row) and Radix natalensis (bottom row). a–d P. columella conchology (a, b), shell microsculpture of the black square hatched area (c) and radular teeth (d) e–h R. natalensis conchology (e, f), shell microsculpture of the black square hatched area (g) and radular teeth (h). Although there is minor variation in the shape of the inner cusp of the first lateral teeth, the discriminatory feature is the periostracum’s spiral ridges