Table 1 Results of generalized linear mixed models (GLMM, logit-link, binomial assumption), and subsequent pairwise comparisons between seasons, respectively, for Haemaphysalis sp. “microcebi”, Lemurpediculus sp., Trombiculidae gen. sp. and Laelaptidae gen. sp.

 Species

-0.537

0.210

1

0.011^*

M. murinus > M. ravelobensis

 Sex

0.278

0.212

1

0.191

Male = female

 Season

4

  EDS vs LRS

1.503

0.738

0.148

  LDS vs LRS

2.715

0.724

<0.001^*

Late dry season > late rainy season

  ERS vs LRS

-0.007

1.233

1.000

  LDS vs EDS

1.212

0.224

<0.001^*

Late dry season > early dry season

  ERS vs EDS

-1.509

1.025

0.412

  ERS vs LDS

-2.721

1.012

0.029^*

Late dry season > early rainy season

 Species

1.446

0.219

1

<0.001^*

M. ravelobensis > M. murinus

 Sex

-0.594

0.382

1

0.120

Male = female

 Season

5

  LDS15 vs EDS15

3.439

0.294

<0.001^*

Late dry season 15 > early dry season 15

  ERS15 vs EDS15

4.963

0.434

<0.001^*

Early rainy season 15 > early dry season 15

  LRS16 vs EDS15

2.897

0.353

<0.001^*

Late rainy season 16 > early dry season 15

  EDS16 vs EDS15

3.84

0.358

<0.001^*

Early dry season 16 > early dry season 15

  ERS15 vs LDS15

1.524

0.331

<0.001^*

Early rainy season 15 > late dry season 15

  LRS16 vs LDS15

-0.542

0.254

0.195

  EDS16 vs LDS15

0.401

0.253

0.489

  LRS16 vs ERS15

-2.067

0.398

<0.001^*

Early rainy season 15 > late rainy season 16

  EDS16 vs ERS15

-1.123

0.395

0.033^*

Early rainy season 15 > early dry season 16

  EDS16 vs LRS16

0.943

0.313

0.020^*

Early dry season 16 > late rainy season 16

Reproductive status males

-1.204

0.277

1

<0.001^*

Testis large > testis small

 Species

-1.375

0.484

1

0.005^*

M. murinus > M. ravelobensis

 Seasonality

3.177

0.655

1

<0.001^*

Dry season > rainy season

 Year

2.264

0.380

1

<0.001^*

2016 > 2015

 Sex

-0.259

0.480

1

0.590

Male = female

 Species

-2.592

0.808

1

0.001^*

M. murinus > M. ravelobensis

 Sex

0.932

0.621

1

0.133

Male = female